After twenty-five years, the authors find that the "natural" writing center-WAC connection has proven its value over time. Pointing to the growth of WAC-writing centers, their scholarship, role in faculty development and community building, the authors end by reflecting on how this program building over the last years supports and can extend our current educational needs for equity, quality online teaching, and community building. [For "The Natural Connection: The WAC Program and the High School Writing Center," see EJ514564.]