From massive open online courses (MOOC) to the smaller scale use of learning management systems, many students interact with online platforms that are meant to support learning. Investigations into the use of these systems have considered how well students learn when certain approaches are employed. However, we do not fully understand how course type or system design influence student actions and experiences, meaning prior findings cannot be properly interpreted and used because we do not understand how these factors influence those findings. Accordingly, we conducted a study to compare student experiences and behaviors across learning management systems and courses from a learning analytics perspective. Differences in student behaviors and experiences highlight how system design and the nature of the course interact: Students reported increased learning support when using a system that foregrounds student interaction through discussion forums, but this relationship did not hold across all course types. Similarly, students from the content-delivery focused system spent more time online while feeling less supported regardless of which type of course they were taking. This newly found evidence for the often-interrelated influence that the course and system have on student experiences and behaviors should therefore be considered when selecting a system to meet particular pedagogical goals.